I found the problem! I forgot to delete the non-deformer history in the model file, which got referenced through the pipeline, and just wrecked havoc all the way down. When I deleted the history (to fix the problem), I lost my blendshape node.
Don't worry though! I reattached the blend shapes, and the animation I made transferred to the new node, no problem. BUT! I used the old blendshape node to drive the teeth and tongue, and all those driven keys were lost. I did a quick repair, and the teeth are looking a little funky, but it's not bad!

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